Dr. Apple received his doctorate degree from the University of Minnesota in 1979 and obtained a fellowship in Laboratory Medicine in 1982 from Washington University. He is board certified in Clinical Chemistry and Toxicology. He serves as the principal investigator of the CLIA-accredited Cardiac Biomarkers Trials Laboratory of the Hennepin Healthcare Research Institute. In addition to his investigative research, Dr. Apple is a Professor at the University of Minnesota Department of Laboratory Medicine and Pathology. Dr. Apple also serves as an Associate Editor for the journal Clinical Chemistry and is a Fellow of the American College of Sports Medicine.