Alumni Directory

Since program inception, 183 trainees have completed MD-PhD training.  A total of 140 (76%) hold research positions in academic medicine or industry, or are still in academic medicine training. In a recent alumni survey, 82% of respondents indicated that they would choose to training again in an MD/PhD program, and 95% of respondents indicated that they were satisfied with the direction of their careers

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Terrence J. Adam

Thesis title: A cost analysis of home montoring in post-lung transplantation patients
Advisor: Stan Finkelstein, Ph.D.
Graduation year: 2003
Graduate program: Health Informatics
Residency: Internal Medicine, Mayo Clinic, Scottsdale, AZ
Current position: Associate Professor, University of Minnesota

Rehana L. Ahmed-Saucedo

Thesis title: Epidemiological associations of metabolic factors with cancer risk and cancer late effects
Advisor: Graduation year: 2007
Graduate program: Epidemiology
Residency: Dermatology, University of Minnesota
Current position: Assistant Professor of Dermatology, University of Minnesota

Benjamin al-Haddad

Thesis Title: Mapping cancer surveillance in Nigeria
Advisor: Logan Spector, Ph.D.
Graduation year: 2015 
Graduate program: Epidemiology 
Residency: Pediatrics, University of Washington 
Current position: Assistant Professor, University of Minnesota

John S. Albin

Thesis title: Human Immunodeficiency Virus evasion of APOBEC3 restriction factors
Advisor: Reuben Harris, Ph.D.
Graduation year: 2014
Graduate program: MICaB
Residency: Internal Medicine, Massachusetts General Hospital
Current position: Instructor in Medicine. Massachusetts General Hospital

Ron C. Anafi

Thesis title: Airway bi-stability in a model for the constricted lung
Advisor: Theodore Wilson, Ph.D.
Graduation year: 2004
Graduate program: Aerospace Engineering and Mechanics
Residency: Internal Medicine, University of Vermont
Current position: Assistant Professor, University of Pennsylvania

Brian M. Andersen

Thesis Title: The role of oxygen in the immunogenicity of brain tumor cell vaccines
Advisors: Walter Low, Ph.D. and Michael Olin, Ph.D.
Graduation year: 2015 
Graduate program: Neuroscience 
Residency: Neurology, Weill Cornell Medical Center 
Current position: Neuro Oncology Fellow, Massachusetts General Hospital

Robert C. Anderson

Thesis title: Genetic analysis of primordial germ cell migration in the mouse
Advisor: Janet Heasman, Ph.D.
Graduation year: 2001
Graduate program: Biomedical Science
Residency: Dermatology, University of Texas, Dallas
Current position: Physician, St. Paul, MN

Shane S. Anderson

Graduation year: 1996
Graduate program: Pathobiology
Residency: Pathology, University of Washington
Current position: Physician, Seattle, WA

Kristin M. Andruska

Thesis title: Partial loss of Tip60 neurodegeneration in a Spinocerebellar Ataxia Type 1 (SCA1) mouse model
Advisor: Harry Orr, Ph.D.
Graduation year: 2011 
Graduate program: BMBB 
Residency: Neurology, Barnes-Jewish Hospital/Washington University 
Current position: Physician

Mathew G. Angelos

Thesis title: Mechanisms of Early Hemato-endothelial Development from Human Pluripotent Stem Cells
Advisor: Dan Kaplan, M.D., Ph.D.
Graduation year: 2018
Graduate program: MICaB
Residency: Internal Medicine, University of Pennsylvania
Current position: Assistant Professor, University of Colorado School of Medicine

Karen R. Armbrust

Thesis title: Murine models of Spinocerebellar Ataxia Type 5
Advisor: Laura Ranum, Ph.D.
Graduation year: 2011
Graduate program: MCDB&G
Residency: Ophthalmology, University of Minnesota
Current position: Assistant Professor of Ophthalmology, University of Minnesota

Michael B. Armstrong

Thesis title: Regulation of Hepatic Uncoupling Protein-2: implications of its role in energy metabolism
Advisor: Howard Towle, Ph.D.
Graduation year: 2001
Graduate program: BMBB
Residency: Pediatrics, University of Michigan, Ann Arbor
Current position: Assistant Professor, Duke University

William F. Auffermann

Thesis title: Template free statistical methods for fMRI date analysis and their applications in clinical medicine
Advisor: Xiaoping Hu, Ph.D.
Graduation year: 2004
Graduate program: Biomedical Engineering
Residency: Radiology, University of Minnesota 
Current position: Assistant Professor, Emory University

Carlos I. Ayala

Thesis title: Rab-GTPase dependent regulation of autophagy and mTOR signaling in the larval fat body of Drosphila melanogaster.
Advisor: Thomas Neufeld, Ph.D.
Graduation year: 2017
Graduate program: Molecular, Cellular, Developmental Biology and Genetics
Residency: Surgery, Stanford University
Current position: Resident

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Aaron M.T. Barnes

Thesis title: Ultrastructural analysis of the extracellular matrix in early Enterococcus faecalis biofilms
Advisor:Gary Dunny, Ph.D.
Graduation year: 2014
Graduate program: MICaB
Residency: Pathology, University of Minnesota
Current position: EPR Laboratory Unit Supervisor, MN Department of Health

Richard J. Battafarano

Graduation year: 1996
Graduate program: Biomedical Sciences
Residency: Surgery, University of Minnesota 
Current position: Associate Professor, University of Maryland

Joan D. Beckman

Thesis title: Regulation and effects of HO-1 expression
Advisor: Gregory Vercellotti, M.D.
Graduation year: 2012
Graduate program: MICaB 
Residency: Internal Medicine, University of North Carolina 
Current position: Assistant Professor of Medicine, Univeristy of Minnesota

Daniel J. Beisang

Thesis title: Regulation and function of the RNA-binding protein CELF1
Advisor: Paul Bohjanen, M.D., Ph.D.
Graduation year: 2014
Graduate program: MICaB
Residency: Pediatrics, University of Minnesota
Current position: Assistant Professor of Pediatrics, University of Minnesota

Scott J. Benson

Thesis title: Genetic analysis of two novel avian endogenous viruses from distinct families
Advisors: Kathleen Conklin, Ph.D. and David Zarkower, Ph.D.
Graduation year: 2002
Graduate program: BMBB 
Residency: Family Practice, United Hospital 
Current position: Physician, Apple Valley, MN

Charles J. Billington Jr.

Thesis title: Mechanisms and prevention of variable craniofacial defects in twisted gastrulation mutant mice
Advisor: Anna Petryk, M.D.
Graduation year: 2014
Graduate program: MCDB&G
Residency: Pediatrics-Medical Genetics, Children's National Medical Center/NIH
Current position: Assistant Professor of Pediatrics, University of Minnesota

Rachael K. Blackman

Thesis Title: Neural basis of context processing dysfunction in schizophrenia: a monkey model
Advisor: Matthew Chafee, Ph.D.
Graduation year: 2015 
Graduate program: Neuroscience 
Residency: Psychiatry, Brown University 
Current position: Physician, Division of Psychiatry- FDA

Sara Bolivar Wagers

Thesis Title: Enhancing regulatory T cell therapy efficacy for acute GVHD prevention
Advisor: Bruce Blazar, M.D.
Graduation year: 2024 
Graduate program: Microbiology, Immunology and Cancer Biology 
Residency:  Pediatrics (PSTP), Children's Hospital of Philadelphia
Current position: Pediatrics Resident, Children's Hospital of Philadelphia

Dieter D. Brandner

Thesis Title: Neuroligin-3 Modulates Opioid-Evoked Changes in Behavior and Brain Function
Advisors: Patrick Rothwell, Ph.D., Paul Mermelstein, Ph.D.
Graduation year: 2024 
Graduate program: Neuroscience
Residency: Ophthalmology, UCSF
Current position: Ophthalmology Resident, UCSF

Julia A. Brekke-Riedl

Thesis Title: Steps towards understanding cellular therapies for Recessive Dystrophic Epidermolysis Bullosa
Advisors: Jakub Tolar, M.D., Ph.D., Bryce Binstadt, M.D., Ph.D.
Graduation year: 2024 
Graduate program: Microbiology, Immunology and Cancer Biology
Residency: Psychiatry, Brown Univ
Current position: Psychiatry Resident, Brown Univ

Terence C. Burns

Thesis title: Stem cells in ischemic brain injury
Advisors: Catherine Verfaillie, M.D. and Walter Low Ph.D.
Graduation year: 2009
Graduate program: Neuroscience 
Residency: Neurosurgery, Stanford University 
Current position: Associate Professor, Mayo Clinic

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Javier Cabrera-Perez

Thesis title: Examining T-cell Recovery After Sepsis 
Advisors: Thomas Griffin, Ph.D. 
Graduation year: 2018
Graduate program: MICaB 
Residency: Pediatrics (PSTP), Baylor College of Medicine 
Current position: Fellow, Brigham and Women's Hospital

Danita J. Carlson

Graduation year: 1995
Graduate program: Neuroscience 
Residency: Internal Medicine, Dartmouth-Hitchcock Medical Center 
Current position: Physician, Minnetonka, MN

Erik S. Carlson

Thesis title: Effects of Genetic and Dietary Iron Deficiency in the Developing Hippocampus.
Advisors: Michael O'Connor, Ph.D. and Michael Georgieff, M.D.
Graduation year: 2010
Graduate program: Neuroscience 
Residency: Psychiatry, University of Washington 
Current position: Assistant Professor of Psychiatry, University of Washington

Jonathan C.T. Carlson

Thesis title: Towards therapeutic nanoassemblies: chemically induced protein oligomerization.
Advisor: Carston Wagner, Ph.D
Graduation year: 2006
Graduate program: Medicinal Chemistry 
Residency: Internal Medicine, Brigham & Women's Hospital/Harvard 
Current position: Assistant Professor of Medicine, Massachusetts General Hospital

Adam F. Carpenter

Graduation year: 2001
Graduate program: Neuroscience 
Residency: Psychiatry, University of Minnesota 
Current position: Assistant Professor, University of Minnesota

Lyn P. Chapman (Hooke)

Graduation year: 1996
Graduate program: Pharmacology 
Residency: Pediatrics, University of Minnesota 
Current position: Physician, Brooklyn Park, MN

Nathan J. Charles

Thesis title: Membrane and nuclear progesterone receptors activate tumor suppressive signaling mechanisms in cell line models of human ovarian cancer”
Advisor: Carol Lange, Ph.D.
Graduation year: 2012
Graduate program: MICaB 
Residency: Pathology, University of Wisconsin, Madison 
Current position: Physician

Eleanor Y. Chen

Thesis title: Gaining insights into vertebrate vascular development: Characterization of zebrafish morphants identified from a morpholino-based vascular screen.
Advisor: Stephen Ekker, Ph.D.
Graduation year: 2006
Graduate program: MCDB&G 
Residency: Pathology, Brigham & Women's Hospital/Harvard 
Current position: Associate Professor of Pathology, University of Washington

Ian N. Cheong

Thesis title: Non-invasive assessment of neurochemical biomarkers in Amyotrophic Lateral Sclerosia
Advisors: Gulin Oz, Ph.D.
Graduation year: 2020
Graduate program: Neuroscience
Residency: Neurology, Johns Hopkins University 
Current position: Resident

Jason A. Chesney

Thesis title: Definition of peripheral blood fibrocytes as novel antigen presenting cells
Advisor: Kathleen Conkin, Ph.D.
Graduation year: 1998
Graduate program: Biomedical Sciences 
Residency: Internal Medicine, New York Hospital 
Current position: Professor and Director, Brown Cancer Center, University of Louisville

Caitlin B. Conboy

Thesis title: R-spondin 2 drives Wnt signaling and tumorigenesis in multiple tissues
Advisor: David Largaespada, Ph.D.
Graduation year: 2016
Graduate program: MICaB 
Residency: Internal Medicine, Mayo Clinic 
Current position: Hematology/Oncology Fellow, Mayo Clinic

Samuel W. Cramer

Thesis title: Normal and diseased circuitry in the cerebellar and cerebral cortex
Advisor: Timothy Ebner, M.D., Ph.D.
Graduation year: 2016
Graduate program: Neuroscience 
Residency: Neurological Surgery, University of Minnesota 
Current position: Resident

Clifford M. Csizmar

Thesis title: Engineered proteins with tunable avidity for studying and controlling cellular interactions
Advisors: Carston Wagner, Ph.D.
Graduation year: 2020
Graduate program: Medicinal Chemistry
Residency: Internal Medicine - CITP, Mayo Clinic 
Current position: Resident

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Matthew J. Deeths

Thesis title: Costimulation of CD8 positive T lymphocytes
Advisor: Matthew Mescher, Ph.D.
Graduation year: 2000 
Graduate program: MICaB
Residency: Dermatology, University of Colorado 
Current position: Physician, Grand Junction, CO

Christopher C. DeNucci

Thesis title: The role of beta-1 integrin on CD4 T cells
Advisor: Yoji Shimizu, Ph.D.
Graduation year: 2012 
Graduate program: MICaB 
Residency: Diagnostic Radiology, Stanford University 
Current position: Physician

Martin M. Dinges

Thesis title: Studies of TSST-1-induced hypersensitivity to LPS
Advisor: Patrick Schlievert, Ph.D.
Graduation year: 2001 
Graduate program: MICaB 
Residency: Pathology, University of Washington 
Current position: Physician, Albuquerque, NM

Melvin T. Donaldson

Thesis title: Yoga for chronic pain in veterans: a mixed methods study 
Advisors: Erin Krebs, M.P.H. and Rich MacLehose, Ph.D.
Graduation year: 2020
Graduate program: Epidemiology
Residency: Emergency Medicine - PSTP, University of Iowa 
Current position: Fellow, University of Iowa

Alexander J. Dwyer

Thesis title: Hybrid peptide-specific T cells incite islet inflammation and autoimmune diabetes 
Advisor: Brian Fife, Ph.D.
Graduation year: 2024
Graduate program: Microbiology, Immunology and Cancer Biology
Residency: Otolaryngology, Johns Hopkins Hospital 
Current position: Resident

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Craig E. Eckfeldt

Thesis title: Discovery and functional analysis of novel determinants of hematopoietic stem and progenitor cell function
Advisor: Catherine Verfaillie, M.D.
Graduation year: 2007 
Graduate program: MICaB 
Residency: Internal Medicine, University of Minnesota 
Current position: Associate Professor of Medicine, University of Minnesota

Benjamin D. Ehst

Thesis title: Design and characterization of a model of murine skin graft rejection: visualizing the immune response against a defined minor transplantation antigen
Advisor: Marc Jenkins, Ph.D.
Graduation year: 2003 
Graduate program: MICaB 
Residency: Dermatology, Johns Hopkins Hospital 
Current position: Assistant Professor, Oregon Health & Science University

Bart T. Endrizzi

Thesis title: The regulation and function of KLF-2 in T cell biology
Advisor: Stephen Jameson, Ph.D.
Graduation year: 2005 
Graduate program: MICaB 
Residency: Dermatology, University of Minnesota 
Current position: Clinical Assistant Professor, University of Minnesota

Kevin C. Engel

Thesis title: Ocular motor and manual tracking of visual targets
Advisor: John Soechting, Ph.D.
Graduation year: 2003 
Graduate program: Neuroscience 
Residency: Ophthalmology, University of Minnesota 
Current position: Chief of Ophthalmology, Hennepin County Medical Center, Minneapolis, MN

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James M. Fink

Graduation year: 1992 
Graduate program: Microbiology 
Residency: Cytogenetics, University of Minnesota
Current position: Assistant Professor, University of Minnesota

Jason D. Fowler

Thesis title: The role of renin in the adipose tissue renin-angiotensin system
Advisors: David Bernlohr, Ph.D. and Steven Katz, Ph.D.
Graduation year: 2011 
Graduate program: Cellular and Integrative Physiology 
Residency: Surgery, University of Minnesota 
Current position: Physician

Daniel T. Franc

Thesis title: Investigations in Magnetic Resonance Diffusion Tensor Imaging applied to clinical and research medicine
Advisor: Kelvin Lim, M.D.
Graduation year: 2010 
Graduate program: Neuroscience 
Residency: Neurology, UCLA 
Current position: Physician

Brigitte L. Frohnert

Thesis title: Functional and regulatory studies of the Fatty Acid Transport Protein 1
Advisor: David Bernlohr, Ph.D.
Graduation year: 2002 
Graduate program: BMBB 
Residency: Pediatrics, University of Minnesota 
Current position: Associate Professor, University of Colorado

Sarah A. Frommer

Thesis title: Investigating the use of multipotent adult progenitor cells for treatment of Duchenne Muscular Dystrophy: a translational approach
Advisors: Catherine Verfaillie, M.D. and Paul Iaizzo, Ph.D.
Graduation year: 2010 
Graduate program: Biomedical Engineering 
Residency: Plastic Surgery, Brown University 
Current position: Physician

Joseph M. Fullmer

Thesis title: Identification of the target glycoproteins of the lectin IB4
Advisor: Robert Elde, Ph.D.
Graduation year: 2006 
Graduate program: Neuroscience 
Residency: Pathology, University of Washington 
Current position: Director of Autopsy Oakland University William Beaumont School of Medicine

Regan S. Fulton

Thesis title: The developmental activation of the kappa immunoglobulin light chain locus
Advisor: Brian Van Ness, Ph.D.
Graduation year: 1996 
Graduate program: Pathobiology 
Residency: Pathology, Stanford University 
Current position: Physician, San Francisco, CA

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Jose E. Gomez Uranga

Thesis title: Reach to grasp representations in the monkey dorsal premotor and primary ,otor cortices: endpoint accuracy and grasp posture
Advisor: Timothy Ebner, M.D., Ph.D.
Graduation year: 2002 
Graduate program: Neuroscience 
Residency: Pediatrics, University of Minnesota 
Current position: Physician, Naval Medical Center, San Diego, CA

Karen S. Gustafson

Thesis title: Transcriptional regulation of HLA-E gene expression by interferon-γ
Advisor: Gordon Ginder, M.D.
Graduation year: 1998 
Graduate program: Biomedical Sciences 
Residency: Pathology, University of Pittsburgh 
Current position: Physician, Philadelphia, PA

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Matthew C. Hagen

Thesis title: Neuroimaging of higher somatosensory processing in humans: Attention, space and motion.
Advisor: Jose Pardo, M.D., Ph.D.
Graduation year: 2005 
Graduate program: Neuroscience 
Residency: Pathology, Indiana University 
Current position: Associate Professor, University of Cincinnati

Michelle Y. Hamline

Thesis title: The best laid plans of mice and men often go awry: roles of BCOR in mammalian embryonic development
Advisor: Vivian Bardwell, Ph.D.
Graduation year: 2013
Graduate program: MCDB&G
Residency: Pediatrics, University of California-Davis
Current position: Assistant Professor of Clinical Pediatrics, University of California, Davis

Maureen Handoko

Thesis title: Specific amyloid-β oligomers in human cerebrospinal fluid
Advisor: Karen Hsiao Ashe, M.D., Ph.D.
Graduation year: 2014
Graduate program: Neuroscience
Residency: Pediatric Neurology, Baylor College of Medicine
Current position: Staff Neurologist, Rocky Mountain Neurodiagnostics

Lauren A. Harasymiw

Thesis title: Centromere mechanical maturation and mitotic fidelity during mammalian cell mitosis
Advisor: Melissa Gardner, Ph.D.
Graduation year: 2019 
Graduate program: BICB 
Residency: Pediatrics (PSTP), University of Minnesota 
Current position: Resident

Jason M. Hataye

Thesis title: Cell competition for survival and proliferation resources defines the CD4+ T cell niche
Advisor: Marc Jenkins, Ph.D.
Graduation year: 2008 
Graduate program: MICaB 
Residency: Internal Medicine, UCLA 
Current position: Physician Scientist, Vaccine Research Center, NIAID

Alan R. Hauser

Graduation year: 1992 
Graduate program: Microbiology
Residency: Infectious Disease, UCSF 
Current position: Professor, Vice Chair, MSTP Director, Northwestern University

Angela L. Hewitt

Thesis Title: Does cerebellar cortex function as a forward internal model for motor control?
Advisor: Timothy Ebner, M.D., Ph.D.
Graduation year: 2015 
Graduate program: Neuroscience 
Residency: Child Neurology, Mayo Clinic 
Current position: Assistant Professor, University of Rochester

Nathan E. Hoffmann

Thesis title: Quantification of host-mediated cryoinjury in urologic tissues
Advisor: John Bischof, Ph.D.
Graduation year: 2003 
Graduate program: Biomedical Engineering 
Residency: Urology, Mayo Clinic, Rochester MN 
Current position: Physician, Duluth, MN

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John R. Jalas

Thesis title: Stereo- and Regioselectivity in the Cytochrome P450-mediate Metabolism of the Tobacco-specific Nitrosamine 4-(Methylnitrosamino)-1-(3-pyridyl)-1-butanone (NNK).
Advisor: Steven Hecht, Ph.D.
Graduation year: 2005 
Graduate program: Chemistry 
Residency: Pathology, UCSF 
Deceased

Tanner M. Johanns

Thesis title: CD4 T cells in protection and pathogenesis of persistent Salmonella infection
Advisor: SingSing Way, M.D., Ph.D.
Graduation year: 2012 
Graduation program: MICaB 
Residency: Internal Medicine, Barnes-Jewish Hospital/Washington University 
Current position: Assistant Professor of Medicine, Washington University

Hemant P. Joshi

Thesis title: Non-canonical pathways of HIF1-alpha regulation in ovarian cancer: implications for angiogenesis and metastasis.
Advisor: Sundaram Ramaskrishnan, Ph.D.
Graduation year: 2013
Graduation program: Pharmacology
Residency: Anesthesiology, University of Florida/Shands Hospital
Current position: Physician

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Kristen M. Kallestad

Thesis title: Duchenne Muscular Dystrophy and extraocular muscle: a potential sparing mechanism with therapeutic implications
Advisor: Linda McLoon, Ph.D.
Graduation year: 2011
Graduate program: Neuroscience 
Residency: Family Medicine, University of Minnesota 
Current position: Physician

Brad S. Karon

Graduation year: 1996 
Graduate program: Biochemistry 
Residency: Pathology, Barnes-Jewish Hospital/Washington University 
Current position: Professory and Dean, Mayo Clinic School of Health Sciences

Sakeen W. Kashem

Thesis title: Immunity against Candida albican skin infections 
Advisor: Dan Kaplan, M.D., Ph.D. 
Graduation year: 2018 
Graduate program: MICaB 
Residency: Dermatology, University of California, San Francisco. Transition year Univ of TX, Southwestern
Current position: Assistant Professor, UCSF

James T. Katter

Graduation year: 1995 
Graduate program: Neuroscience 
Residency: Medicine/Pediatrics, Brigham & Women's Hospital/Harvard 
Current position: Physician, Zimmerman, MN

Christopher Dirk Keene

Thesis title: Experimental Huntington’s Disease – neuroprotective effects of Tauroursodeoxycholic Acid and 1H NMR spectroscopic characterizaion
Advisor: Walter Low, Ph.D.
Graduation year: 2005 
Graduate program: Neuroscience 
Residency: Pathology, University of Washington 
Current position: Professor, University of Washington

Hyon J. Kim

Thesis title: Genetic analysis of early vertebrate endocrine pancreas formation
Advisor: Stephen Ekker, Ph.D.
Graduation year: 2007 
Graduate program: MCDB&G
Residency: Internal Medicine, University of Cincinnati 
Current position: Physician

Yong Y. Kim

Thesis title: Two barriers to RAS mediated oncogenesis: translational control checkpoint and proliferative block by autophagy
Advisor: Peter Bitterman, M.D.
Graduation year: 2011 
Graduate program: MICaB 
Residency: Pediatrics, University of Florida 
Current position: Hematology/Oncology Fellow, Children's National Medical Center

Ivan A. Klement

Thesis title: The role of nuclear localization of Ataxin-1 in the aathogenesis of Spinocerebellar Ataxia Type 1
Advisor: Harry Orr, Ph.D.
Graduation year: 2000 
Graduate program: MICaB 
Residency: Pathology, UCLA 
Current position: Physician, Santa Barbara, CA

LuAnne M. Klemme

Thesis title: Molecular genetic characterization and cloning of the gene coding for the rat acidic epididymal glycoprotein D
Advisor:David Hamilton, Ph.D.
Graduation year: 1997 
Graduate program: Biomedical Sciences 
Residency: Medicine/Pediatrics, University of Hawaii 
Deceased

David A. Knorr

Thesis title: Engineering human pluripotent stem cells for enhanced lymphocyte development and function
Advisor: Dan Kaufman, M.D., Ph.D.
Graduation year: 2014
Graduate program: MICaB
Residency: Internal Medicine PSTP, Weill Cornell Medical Center
Current position: Medical Director, Clinical Development - Oncology and Translational Sciences, Regeneron

Lisa K. Koch

Thesis title: Indoleamine 2, 3-dioxygenase: a potent regulator of graft versus host disease with potential therapeutic applications
Advisor: Bruce Blazar, M.D.
Graduation year: 2011 
Graduate program: MICaB 
Residency: Pathology, University of Washington 
Current position: Acting Assistant Professor of Pathology, University of Washington

Marcos T. Kuroki

Thesis Title: Neural control of the splanchnic circulation in AngII-salt hypertension
Advisor: John Osborn, Ph.D.
Graduation year: 2015 
Graduate program: Neuroscience 
Residency: Surgery, Pennsylvania State College of Medicine 
Current position: Clinical Fellow, Stanford University

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Christine S. Lambert

Thesis Title: Occupational exposures and the development of lung disease in Minnesota miners
Advisor: Bruce Alexander, Ph.D.
Graduation year: 2015 
Graduate program: Environmental Health Sciences 
Residency: Internal Medicine, University of Minnesota 
Current position: Assistant Professor, University of Minnesota

Michael D. Lane

Thesis title: Developing methods to understand and engineer protease cleavage specificity
Advisors: Burkhard Seelig, Ph.D.
Graduation year: 2018
Graduate program: BMBB

Frederick J.P. Langheim

Thesis title: Dynamic brain interactions as revealed by Magnetoencephalography (MEG).
Advisors: Apostolos Georgopoulos, Ph.D and Arthur Leuthold, Ph.D.
Graduation year: 2007 
Graduate program: Neuroscience 
Residency: Psychiatry, University of Wisconsin 
Current position: Clinical Adjunt Assistant Professor of Psychiatry, University of Wisconsin

Andrew W. Larson

Graduation year: 1995 
Graduate program: Pathobiology 
Residency: Pediatrics, University of Minnesota 
Current position: Physician, Maple Grove, MN

Tuong H. Le

Graduation year: 1998 
Graduate program: Biomedical Engineering 
Residency: Radiology, UCSF 
Current position: Physician, Plano, TX

Joseph Lee

Thesis title: Modulations of myocardial energy metabolism via xathine oxidase inhibition, SCA-1+ stem cell treatment and PPAR-δ gene ablation
Advisor: Jianji Zhang, M.D., Ph.D.
Graduation year: 2008 
Graduate program: Biomedical Engineering
Residency: Radiation Oncology, Barnes-Jewish Hospital/Washington University
Current position: Physician

Peter K. Lee

Thesis title: Studies of the mechanism of hypotension in Toxic Shock syndromes
Advisor: Patrick Schlievert, Ph.D.
Graduation year: 1993 
Graduate program: Microbiology 
Residency: Dermatology, Brigham & Women's Hospital/Harvard 
Current position: Associate Professor, University of Minnesota

Robin D. Lee

Thesis title: Transcriptional and epigenetic networks governing B cell development and transformation
Advisor: Michael Farar, Ph.D.
Graduation year: 2024 
Graduate program: Microbiology, Immunology and Cancer Biology 
Residency: Internal Medicine (PSTP), UMN 
Current position: Resident

Ava Y. Lin

Thesis title: From structure & dynamics to therapeutic developments for neuromuscular disease
Advisor: David Thomas, Ph.D.
Graduation year: 2014
Graduate program: BMBB
Residency: Pediatric Neurology, University of Washington
Current position: Assistant Professor of Pediatric Neurology, University of Michigan

Hong-Yiou Lin

Thesis title: The role of PKC in differential morphine tolerance
Advisor: Horace Loh, Ph.D.
Graduation year: 2014
Graduate program: Pharmacology
Residency: Radiation Oncology, Beaumont Health System
Current position: Physician

Michael A. Linden

Thesis title: Novel targeted deregulation of apoptotic and oncogenic pathways leads to B-lymphoid malignancies in genetically engineered mice
Advisor: Brian Van Ness, Ph.D.
Graduation year: 2006 
Graduate program: MDCB&G 
Residency: Pathology, University of Washington 
Current position: Professor, University of Minnesota

James F. List

Thesis title: Visna virus replication: provirus formation and integration
Advisor: Ashley Haase, M.D.
Graduation year: 1996 
Graduate program: Microbiology 
Residency: Medicine/Pediatrics, Brigham & Women's Hospital/Harvard 
Current position: Vice President, Full Development, Bristol-Myers Squibb

Jennifer L. Long

Thesis title: Elastogenesis in cardiovascular tissue engineering
Advisor: Robert Tranquillo, Ph.D.
Graduation year: 2004
Graduate program: Chemical Engineering 
Residency: Otolaryngology, UCLA 
Current position: Associate Professor, UCLA

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Shawn A. Mahmud

Thesis title: TNF receptor superfamily costimulation drives thymic Treg development and repertoire selection
Advisor: Mike Farrar, Ph.D.
Graduation year: 2016 
Graduate program: MICaB
Residency: Pediatrics, University of Minnesota 
Current position: Pediatric Rheumatology Fellow, University of Minnesota

Erika-Nell Malvey-Dorn

Thesis title: The functional consequences of T cell tolerance induction in vivo
Advisor: Daniel Mueller, M.D.
Graduation year: 2000 
Graduate program: MICaB 
Residency: Pediatrics, University of Minnesota 
Current position: Physician, Eden Prairie, MN

Tracy A. Marko (Powell)

Thesis title: Uncovering Mechanisms of Osteosarcoma Metastasis
Advisor: David Largaespada, Ph.D.
Graduation year: 2018 
Graduate program: MICaB 
Residency: Emergency Medicine, Regions Hospital, St. Paul, MN 
Current position: Resident

Stacene R. Maroushek

Thesis title: Sequence analysis of a viral gene found within the central nervous system of patients with human T-cell Lymphotropic Virus Type 1-Associated Disease
Advisor: Ashley Haase, M.D.
Graduation year: 1994 
Graduate program: Microbiology 
Residency: Pediatrics, University of Minnesota 
Current position: Assistant Professor, University of Minnesota

Cameron McDonald-Hyman

Thesis title: Methods to enhance the efficacy of Regulatory T-cell therapies for acute and chronic Graft-versus-Host Disease
Advisor: Bruce Blazar, M.D.
Graduation year: 2018 
Graduate program:  MICaB
Residency: Internal Medicine, University of Michican, Ann Abor
Current position: Fellow, University of Minnesota

Benjamin Mueller

Thesis title: Direct detection of phospholamban and SERCA interaction in membranes using fluorescence resonance energy transfer
Advisor: David Thomas, Ph.D.
Graduation year: 2006 
Graduate program: BMBB 
Residency: Orthopaedic Surgery, University of Minnesota 
Current position: Physician, Minneapolis, MN

Debra L. Murray

Thesis title: Structure-function studies of Toxic Shock Syndrome Toxin-1
Advisor: Patrick Schlievert, Ph.D.
Graduation year: 1997 
Graduate program: Microbiology 
Residency: Internal Medicine, Vanderbilt University 
Current position: Physician, Tulsa, OK

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Jennifer C. Nagode

Thesis title: Neuroimaging Studies of the Medial Temporal Lobe Role During Learning and Memory
Advisor: Jose Pardo, M.D., Ph.D.
Graduation year: 2004 
Graduate program: Neuroscience 
Residency: Psychiatry, UCSD 
Current position: Physician, San Diego, CA

Minelva R. Nanton

Thesis Title: Protective B cell responses during Salmonella infection
Advisors: Stephen McSorley, Ph.D. and Chandy John, M.D.
Graduation year: 2015 
Graduate program: MICaB 
Residency: Family Medicine, University of Minnesota 
Current position: Physician

Ryan W. Nelson

Thesis Title: The origin and function of a large naïve CD4+ T cell population
Advisors: Marc Jenkins, Ph.D.
Graduation year: 2016 
Graduate program: MICaB 
Residency: Pediatrics, Boston Children's Hospital 
Current position: Assistant Professor, University of Minnesota

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Scott A. Oakman

Graduation year: 1999 
Graduation program: Neuroscience 
Residency: Psychiatry, University of Michigan 
Current position: Assistant Professor, University of Minnesota

Oludare Odumade

Thesis title: Prospective analysis of the primary immune response during natural Epstein-Barr virus infection in humans.
Advisor: Kristin Hogquist, Ph.D.
Graduation year: 2014
Graduation program: MICaB
Residency: Pediatrics, University of California-San Diego
Current position: Staff Physician/Instructor, Boston Children's Hospital/Harvard University

Timothy H. Olson

Thesis title: The Acid-Sensing Ion Channels (ASICs): Distribution within Primary Afferent Neurons and Potential Role in Hyperalgesia.
Advisor: Robert Elde, Ph.D.
Graduation year: 2002 
Graduation program: Neuroscience 
Residency: Anesthesiology, Massachusetts General Hospital 
Current position: Physician, St. Cloud, MN

Xilma R. Ortiz-Gonzalez

Thesis title: In vivo neutral differentiation potential of bone marrow-derived multipotent adult progenitor cells.
Advisor: Walter Low, Ph.D.
Graduation year: 2006 
Graduation program: Neuroscience 
Residency: Pediatric Neurology, Children's Hospital of Philadelphia 
Current position: Assistant Professor of Nuerology, Children's Hospital of Philadelphia

Matthew J. O'Shaughnessy

Thesis title: Novel Molecular Strategies for Prevention of Graft-Versus-Host Disease by Ex Vivo Tolerance Induction.
Advisor: Bruce Blazar, M.D.
Graduation year: 2008 
Graduation program: MICaB 
Residency: Urology, University of Minnesota 
Current position: Physician

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Judit M. Perez Ortiz

Thesis title: PKA phosphorylation of ATAXIN1 in Purkinje cells modulates early onset of ataxia
Advisor: Harry Orr, Ph.D.
Graduation year: 2018 
Graduate program: Neuroscience 
Residency: Child Neurology, Mayo Clinic
Current position: Resident

Mark A. Pirner

Graduation year: 1996 
Graduate program: Biomedical Sciences 
Residency: Medicine, Duke University 
Current position: Senior Director and Head, PepsiCo Biology Innovation Laboratory, Purchase, NY

Alexa A. Pragman

Thesis title: Studies on the Staphylococcal Respiratory Response (SrrAB) and its role in Staphylococcus aureus virulence factor regulation
Advisor: Patrick Schlievert, Ph.D.
Graduation year: 2007 
Graduate program: MICaB 
Residency: Internal Medicine, University of Chicago 
Current position: Associate Professor, University of Minnesota

Jonathan T. Pribila

Thesis title: Molecular mechanisms for the recruitment and retention of leukocytes in lymph nodes
Advisor: Yoji Shimizu, Ph.D.
Graduation year: 2005 
Graduate program: MICaB 
Residency: Ophthalmology, University of Michigan 
Current position: Physician, St. Louis Park, MN

Kurt W. Prins

Thesis title: Analysis of cytoskeletal interactions and their implications in skeletal muscle disease
Advisor: James Ervasti, Ph.D.
Graduation year: 2012 
Graduate program: BMBB 
Residency: Internal Medicine, University of Alabama-Birmingham 
Current position: Associate Professor, University of Minnesota

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Erik A. Ranheim

Graduation year: 1996 
Graduation program: Biomedical Sciences 
Residency: Pathology, Stanford University 
Current position: Professor and Chair of Pathology and Laboratory Medicine, University of Wisconsin

Morayma G. Reyes

Thesis title: Purification, Characterization, and Differentiation of Multipotent Adult Stem Cell From Post-Natal Human Bone Marrow.
Advisor: Catherine Verfaillie, M.D.
Graduation year: 2003 
Graduation program: MICaB 
Residency: Clinical Pathology, University of Washington 
Current position: Medical Director of Thrombosis and Hemostasis Labs, Cleveland Clinic

Stephanie Y. Rhee

Thesis title: Regulation of T follicular helper cells in the germinal center reaction of chronic graft-versus-host disease
Advisor: Bruce Blazar, M.D.
Graduation year: 2024 
Graduation program: MICaB 
Residency: Diagnostic Radiology, Mayo Clinic 
Current position: Resident

Benjamin I. Rosner

Thesis title: Contact Guiding Neurotrophic Collagen Gel Rods for Peripheral Nerve Regeneration.
Advisor: Robert Tranquillo, Ph.D.
Graduation year: 2003 
Graduation program: Chemical Engineering 
Residency: Internal Medicine, Kaiser Permanente, Santa Clara, CA 
Current position: Resident

Jared H. Rowe

Thesis title: Regulatory T cells and the immune pathogenesis of prenatal Listeriosis.
Advisor: SingSing Way, M.D., Ph.D.
Graduation year: 2014
Graduation program: MICaB
Residency: Pediatrics, Boston Children's Hospital
Current position: Clinical Pediatric Fellow, Boston Children's Hospital

John N. Rydberg

Graduation year: 1996
Graduation program: Biomedical Engineering 
Residency: Radiology, Mayo Clinic 
Current position: Assistant Program Director, Nuclear Medicine, Oakland University William Beaumont School of Medicine

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José E. San Miguel-Ruiz

Thesis title: The role of the actin-binding protein Arp2/3 in growth coe actin dynamics and guidance
Advisor: Paul Letourneau, Ph.D.
Graduation year: 2016
Graduate program: Neuroscience
Residency: Orthopaedic Surgery, University of Minnesota
Current position: Resident

Jason M. Schenkel

Thesis title: The distribution and function of resident memory CD8 T-cells
Advisor: David Masopust, Ph.D.
Graduation year: 2016
Graduate program: MICaB
Residency: Clincal Pathology, Brigham & Women's Hospital
Current position: Assistant Professor, MD Anderson Cancer Center

Stephen C. Schmechel

Graduation year: 1998 
Graduate program: Microbiology 
Residency: Pathology, University of Washington 
Current position: Associate Professor, University of Washington

Megan M. Schmit

Thesis title: The elucidation of MCM10's role in human disease
Advisor: Anja Bielinsky, Ph.D.
Graduation year: 2024
Graduate program: BMBB
Residency: Pediatrics (PSTP), UMN
Current position: Resident

Kristin L. Schreiber

Thesis title: Neural-Immune Interactions: Studies of the Reciprocal Influence of the Nervous and Immune Systems in Infection and Pain.
Advisors: George Wilcox, Ph.D. and David Brown, Ph.D.
Graduation year: 2007 
Graduate program: Neuroscience 
Residency: Anesthesiology, University of Pittsburgh 
Current position: Associate Professor, Brigham & Women's Hospital/Harvard University

Wade L. Schulz

Thesis title: Cellular Determinants of Reovirus Entry and Infection.
Advisor: Leslie Schiff, Ph.D.
Graduation year: 2014
Graduate program: MICaB
Residency: Clinical Pathology, Yale University
Current position: Assistant Professor of Lab Medicine, Yale University

Robert E. Schwartz

Graduation year: 2006 
Graduate program: Biomedical Engineering 
Residency: Internal Medicine, Barnes-Jewish Hospital/Washington University 
Current position: Associate Professor, Weill Cornell School of Medicine

Mithun M. Shenoi

Thesis title: Nanoparticle preconditioning for enhanced thermal therapies in cancer.
Advisor: John Bischof, Ph.D.
Graduation year: 2013
Graduate program: Biomedical Engineering
Residency: General Surgery, Duke University
Current position: Colorectal Surgery Fellow, Indiana University

Javier E. Sierra-Pagan

Thesis title: Epigenetic regulatory mechanisms that govern cardiovascular development
Advisor: Daniel Garry, M.D.
Graduation year: 2024
Graduate program: MCDBG
Residency: Internal Medicine (PSTP), Univ of Pennsylvania
Current position: Resident

Amila Silva

Thesis title: Multiple roles of Delta-Notch signaling in retinal development.
Advisor: Steven McLoon, Ph.D.
Graduation year: 2001 
Graduate program: Neuroscience 
Residency: Ophthalmology, Mayo Clinic 
Current position: Physician, Fontana, CA

Nicole E.B. Skinner

Thesis Title: Strategies to improve gene expression and targeting for DNA vaccine development
Advisor: Christopher Pennell, Ph.D.
Graduation year: 2015 
Graduate program: MICaB 
Residency: Internal Medicine, Beth Israel Deaconess Medical Center 
Current position: Assistant Professor, Ohio State University College of Medicine

Iberia Romina Sosa

Thesis title: Alpha-BTX-Sensitive nAChRs Modulate Human Bronchial Epithelial Cell Functions.
Advisor: Bianca Conti-Fine, M.D.
Graduation year: 2007 
Graduate program: Neuroscience 
Residency: Internal Medicine, Vanderbilt University 
Current position: Associate Professor and Chief of Hematology, Fox Chase Cancer Center

Anja I. Srienc

Thesis title: Retinal Ischemia in the In Vivo Rodent: Cellular and Vascular Characteristics.
Advisor: Eric Newman, Ph.D.
Graduation year: 2017 
Graduate program: Neuroscience 
Residency: Neurological Surgery, Washington University/Barnes-Jewish Hospital 
Current position: Resident

Jack H. Staddon

Thesis title: Identification and Genetic Characterization of a Group II Intron Insertion Site and Origin of Transfer Region in the Enterococcus faecalis plasmid, pCF10.
Advisor: Garry Dunny, Ph.D.
Graduation year: 2007
Graduate program: MICaB 
Residency: Pediatrics, Loma Linda University 
Current position: Assistant Professor, University of Kansas

Eric R. Stevens

Thesis title: How D-serine and glycine control NMDA receptors in the vertebrate retina.
Advisor: Robert Miller, M.D.
Graduation year: 2007
Graduate program: Neuroscience
Residency: Ophthalmology, Indiana University
Current position: Physician

Jared D. Sturgeon

Thesis title: Risk factors for intraparenchymal hemorrhagic stroke and Twin-Cities area stroke trends
Advisor: Aaron Folsom, M.D., M.P.H.
Graduation year: 2008
Graduate program: Epidemiology
Residency: Radiation Oncology, University of Texas-Galveston
Current position: Physician

Brian M. Sweis

Thesis title: Beyond simple tests of value: a neuroeconomic, translational, disease-relevant, and circuit-based approach to resolve the computational complexity of decision making
Advisors: Mark Thomas, Ph.D. and David Redish, Ph.D.
Graduation year: 2020
Graduate program: Neuroscience
Residency: Psychiatry - PSTP, Icahn School of Medicine/Mount Sinai 
Current position: Assistant Professor, Icahn School of Medicine/Mount Sinai

Thomas M. Suszynski

Thesis title: Tissue-engineered islet graft design and assessment
Advisor: Klearchos Papas, Ph.D.
Graduation year: 2014
Graduate program: Biomedical Engineering
Residency: Plastic Surgery, University of Texas-Southwestern
Current position: Assistant Professor, Dept of Surgery, University of Minnesota

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Annie Tan

Thesis title: The role of cyclin D1 in survival signaling downstream of Eukaryotic Translation Initiation Factor 4E
Advisors: Peter Bitterman, M.D. and James McCarthy, Ph.D.
Graduation year: 2000 
Graduate program: MICaB 
Residency: OB/GYN, UCSF 
Current position: Physician, Minneapolis, MN

Randall S. Taylor

Thesis title: Proteomic studies of the rat liver golgi complex: calcium uptake, 2D family gel analysis, protein identification and characterization of the RLP30 family of proteins
Advisor: Kathryn Howell, Ph.D.
Graduation year: 2001 
Graduate program: Cell and Developmental Biology, University of Colorado 
Residency: ENT, University of Minnesota 
Current position: Physician, Littleton, CO

David G. Telander

Graduation year: 1999 
Graduateprogram: MIMP 
Residency: Ophthalmology, University of Minnesota 
Current position: Physician, Sacramento, CA

Christina M.B.T. Thaler

Thesis Title: Modulation of BMP signaling during dorsal ventral patterning in Drosophila melanogaster
Advisors: Michael O'Connor, Ph.D. and Hans Othmer, Ph.D.
Graduation year: 2015 
Graduate program: Chemical Engineering 
Residency: Internal Medicine, Abbott Northwestern Hospital 
Current position: Cardiovascular Disease Fellow, Hennepin County Medical Center

Serena K.A. Thompson

Thesis title: Temporal and spatial properties of the fMRI response to first and second order contrast in V1
Advisor: Daniel Kersten, Ph.D.
Graduation year: 2012 
Graduate program: Neuroscience 
Residency: Neurology, Medical College of Wisconsin 
Current position: Assistant Professor of Neurology, Medical College of Wisconsin

Philip J. Titcombe

Thesis title: Epigenetic regulation of T cell peripheral tolerance by a BATF-BIM axis
Advisor: Daniel Mueller, M.D.
Graduation year: 2024 
Graduate program: MICaB 
Residency: Internal Medicine (PSTP), Yale 
Current position: Resident

Elizabeth D. Trice Loggers

Thesis title: The role of medical futility in assignment of DNR orders among hospitalized stroke patients
Advisor: Rosalie Kane, Ph.D.
Graduation year: 2003 
Graduate program: Health Services Research 
Residency: Internal Medicine, Massachusetts General Hospital 
Current position: Assistant Investigator, Group Health Research Institute, Seattle, WA

Brian H. Trieu

Thesis title: A non-canonical role of angiotensin-converting enzyme in synaptic plasticity
Advisor: Patrick Rothwell, Ph.D.
Graduation year: 2024 
Graduate program: Neuroscience 
Residency: Psychiatry, Mayo Clinic 
Current position: Resident

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Kevin B. Urdahl

Graduation year: 1995 
Graduation program: Microbiology 
Residency: Pediatrics, University of Washington 
Current position: Professor, Seattle Children's Research Institute

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Ruben A. Velazquez

Thesis title: Effect of capsaicin on zinc localization, Preprotachykinin and Neurokinin-1 messenger Ribonucleic Acid content and Neurokinin-1 receptor internalization: comparison to the effect produced by cubstance P N-Terminal fragments
Advisor: Alice Larson, Ph.D.
Graduation year: 2000 
Graduate program: Neuroscience 
Residency: ENT, University of Texas-San Antonio 
Current position: Physician, Mayaguez, PR

Mayank Verma

Thesis title: Cross talk between skeletal muscle stem cell and endothelial stem cell
Advisors: Atsushi Asakura, Ph.D.
Graduation year: 2020
Graduate program: Integrative Biology and Physiology
Residency: Child Neurology/Neuroscience, University of Texas - Southwestern 
Current position: Resident

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Kirk J. Wangensteen

Thesis title: Expression of transgenes in liver by gene therapy-mediated repopulation
Advisor: Stephen Ekker, Ph.D.
Graduation year: 2009 
Graduate program: BMBB 
Residency: Internal Medicine, University of Pennsylvania
Current position: Associate Professor, Mayo Clinic

Kelsey M. Wanhainen

Thesis title: P2RX7: a double-edged sword in tumor immunotherapy
Advisor: Stephen Jameson, Ph.D.
Graduation year: 2024 
Graduate program: MICaB 
Residency: Internal Medicine, UCLA
Current position: Resident

Christopher A. Warlick

Thesis title: Selective expansion of cells expressing drug-resistant dihydrofolate reductase activity
Advisor: R. Scott McIvor, Ph.D.
Graduation year: 2001 
Graduate program: Pathobiology 
Residency: Urology, Johns Hopkins University 
Current position: Associate Professor and Head of Urology, University of Minnesota

Scott G. Warren

Thesis title: Selective Attentional Modulation of Task-Appropriate Neural Populations in Primary Visual Cortex
Advisor: Geoff Ghose, Ph.D.
Graduation year: 2018
Graduate program: Neuroscience 
Residency: Neurology, Rhode Island Hospital/Brown Univeristy
Current position: Resident

Matthew Wheelwright

Thesis title: iPSC-Derived Cardiomyocytes: Towards an in vitro Model of Cardiac Physiology
Advisor: Joseph Metzger, Ph.D.
Graduation year: 2019
Graduate program: Integrative Biology and Physiology
Residency: Surgery, Univeristy of Minnesota
Current position: Resident

Alexa R. Weingarden

Thesis title: Mechanisms of Fecal Microbiota Transplantation and Development of Novel Therapies for Clostridium difficile Infection.
Advisors: Michael Sadowsky, Ph.D. and Alexander Khoruts, M.D.
Graduation year: 2017 
Graduate program: MICaB 
Residency: Internal Medicine, Stanford University 
Current position: Assistant Professor, University of Minnesota

Michael A. Weinreich

Thesis title: The K, the L, the F, the 2, and the ology
Advisor: Kristin Hogquist, Ph.D.
Graduation year: 2012 
Graduate program: MICaB 
Residency: Pediatrics, University of Maryland 
Current position: VP & Head of Immunology & Inflammation Clinical Development, Apollo Therapeutics

Nathan E. Welty

Thesis title: In vivo functions of intestinal dendritic cells
Advisor: Dan Kaplan, M.D., Ph.D.
Graduation year: 2016
Graduate program: MICaB 
Residency: Internal Medicine, University of Pennsylvania
Current position: Hematology/Oncology Fellow, University of Pennsylvania

Christopher T. Wilke

Thesis title: Causal-based analysis of epileptogenic networks
Advisor: Bin He, Ph.D.
Graduation year: 2011 
Graduate program: Biomedical Engineering 
Residency: Radiation Oncology, University of Minnesota 
Current position: Assistant Professor, University of Pittsburgh

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Amritha Yellamilli

Thesis title: Side population cells in the adult heart
Advisors: Jop van Berlow, Ph.D.
Graduation year: 2020
Graduate program: Integrative Biology and Physiology
Residency: Pediatrics - Research Track, Stanford University
Current position: Resident

Z

Jennifer L. Zick

Thesis title: Effective disconnection of intrinsic networks in the prefrontal cortex: convergence across primate and mouse models of schizophrenia
Advisors: Tay Netoff, Ph.D. and Matt Chafee, Ph.D.
Graduation year: 2020
Graduate program: Neuroscience
Residency: Psychiatry - PSTP, University of Minnesota
Current position: Resident

Jeannette M. Zinggler Berg

Thesis title: Metabolism of nicotine and the tobacco carcinogen 4-(methylnitrosamino)-1-(3-pyridyl)-1-butanone (NNK): genetic and phenotypic variation
Advisor: Sharon Murphy, Ph.D.
Graduation year: 2011 
Graduate program: BMBB 
Residency: Internal Medicine, Vanderbilt University 
Current position: Physician